PUBLICATION

Ripply suppresses Tbx6 to induce dynamic-to-static conversion in somite segmentation

Authors
Yabe, T., Uriu, K., Takada, S.
ID
ZDB-PUB-230414-75
Date
2023
Source
Nature communications   14: 21152115 (Journal)
Registered Authors
Takada, Shinji
Keywords
none
MeSH Terms
  • Animals
  • Gene Expression Regulation, Developmental
  • Mesoderm/metabolism
  • Somites*/metabolism
  • T-Box Domain Proteins/genetics
  • T-Box Domain Proteins/metabolism
  • Zebrafish*/genetics
  • Zebrafish*/metabolism
  • Zebrafish Proteins/genetics
  • Zebrafish Proteins/metabolism
PubMed
37055428 Full text @ Nat. Commun.
Abstract
The metameric pattern of somites is created based on oscillatory expression of clock genes in presomitic mesoderm. However, the mechanism for converting the dynamic oscillation to a static pattern of somites is still unclear. Here, we provide evidence that Ripply/Tbx6 machinery is a key regulator of this conversion. Ripply1/Ripply2-mediated removal of Tbx6 protein defines somite boundary and also leads to cessation of clock gene expression in zebrafish embryos. On the other hand, activation of ripply1/ripply2 mRNA and protein expression is periodically regulated by clock oscillation in conjunction with an Erk signaling gradient. Whereas Ripply protein decreases rapidly in embryos, Ripply-triggered Tbx6 suppression persists long enough to complete somite boundary formation. Mathematical modeling shows that a molecular network based on results of this study can reproduce dynamic-to-static conversion in somitogenesis. Furthermore, simulations with this model suggest that sustained suppression of Tbx6 caused by Ripply is crucial in this conversion.
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