PUBLICATION

Protein tyrosine phosphatase PTPN9 regulates erythroid cell development through STAT3 dephosphorylation in zebrafish

Authors
Bu, Y., Su, F., Wang, X., Gao, H., Lei, L., Chang, N., Wu, Q., Hu, K., Zhu, X., Chang, Z., Meng, K., Xiong, J.W.
ID
ZDB-PUB-140513-152
Date
2014
Source
Journal of Cell Science   127(Pt 12): 2761-70 (Journal)
Registered Authors
Xiong, Jing-Wei
Keywords
none
MeSH Terms
  • Animals
  • Embryo, Nonmammalian/cytology
  • Embryo, Nonmammalian/enzymology
  • Erythroid Cells/enzymology*
  • Erythropoiesis
  • GATA1 Transcription Factor/metabolism
  • Gastrulation
  • Gene Expression
  • Gene Expression Regulation, Developmental
  • Humans
  • K562 Cells
  • Phosphorylation
  • Protein Processing, Post-Translational*
  • Protein Tyrosine Phosphatases, Non-Receptor/physiology*
  • STAT3 Transcription Factor/metabolism*
  • Transcription Factors/metabolism
  • Zebrafish/embryology
  • Zebrafish/physiology*
  • Zebrafish Proteins/metabolism*
  • Zebrafish Proteins/physiology*
PubMed
24727614 Full text @ J. Cell Sci.
Abstract
Protein tyrosine phosphatases (PTPs) such as SHP-1, SHP-2 and CD45 are involved in hematopoiesis, but the function of many PTPs is not well characterized in vivo. Here we have identified Ptpn9a, an ortholog of human PTPN9, as a crucial regulator of erythroid cell development in zebrafish embryos. ptpn9a, but not ptpn9b, was expressed in the posterior lateral plate mesoderm and intermediate cell mass, two primitive hematopoietic sites during zebrafish embryogenesis. Morpholino-mediated knockdown of ptpn9a depleted erythrocytes by inhibiting erythroid cell maturation without affecting erythroid proliferation and apoptosis. Consistently, both dominant-negative PTPN9C515S and PTPN9 siRNA inhibited erythroid differentiation in human K562 cells. Mechanistically, depletion of PTPN9 in zebrafish embryos in vivo or K562 cells in vitro increased phosphorylated STAT3 (pSTAT3), and the hyper-phosphorylated STAT3 entrapped and prevented GATA1 and ZBP-89 from regulating erythroid gene expression. These findings imply that PTPN9 plays an important role in erythropoiesis by disrupting an inhibitory complex of pSTAT3, GATA1 and ZBP-89, providing new cellular and molecular insights of ptpn9a into developmental hematopoiesis.
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