PUBLICATION

Evolution of Nova-Dependent Splicing Regulation in the Brain

Authors
Jelen, N., Ule, J., Ivin, M., and Darnell, R.B.
ID
ZDB-PUB-071023-15
Date
2007
Source
PLoS Genetics   3(10): 1838-1847 (Journal)
Registered Authors
Keywords
Zebrafish, Chickens, Alternative splicing, Mammalian genomics, RNA splicing, Sequence motif analysis, Sequence alignment, Sequence analysis
MeSH Terms
  • Alternative Splicing*
  • Amyloidogenic Proteins/genetics*
  • Animals
  • Antigens, Neoplasm/genetics*
  • Brain/metabolism*
  • Chickens
  • Evolution, Molecular
  • Humans
  • Mice
  • Models, Biological
  • Multigene Family
  • Nerve Tissue Proteins/genetics*
  • Opossums
  • RNA-Binding Proteins/genetics*
  • Species Specificity
  • Xenopus
  • Zebrafish
  • Zebrafish Proteins/genetics*
PubMed
17937501 Full text @ PLoS Genet.
Abstract
A large number of alternative exons are spliced with tissue-specific patterns, but little is known about how such patterns have evolved. Here, we study the conservation of the neuron-specific splicing factors Nova1 and Nova2 and of the alternatively spliced exons they regulate in mouse brain. Whereas Nova RNA binding domains are 94% identical across vertebrate species, Nova-dependent splicing silencer and enhancer elements (YCAY clusters) show much greater divergence, as less than 50% of mouse YCAY clusters are conserved at orthologous positions in the zebrafish genome. To study the relation between the evolution of tissue-specific splicing and YCAY clusters, we compared the brain-specific splicing of Nova-regulated exons in zebrafish, chicken, and mouse. The presence of YCAY clusters in lower vertebrates invariably predicted conservation of brain-specific splicing across species, whereas their absence in lower vertebrates correlated with a loss of alternative splicing. We hypothesize that evolution of Nova-regulated splicing in higher vertebrates proceeds mainly through changes in cis-acting elements, that tissue-specific splicing might in some cases evolve in a single step corresponding to evolution of a YCAY cluster, and that the conservation level of YCAY clusters relates to the functions encoded by the regulated RNAs.
Genes / Markers
Figures
Show all Figures
Expression
Phenotype
Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
Engineered Foreign Genes
Mapping